Richard
I think I might have found a trail for James Lane.
Charles Henry Lane, B 1846 Herne Hill, (son of James Lane and Elizabeth Horsburgh); 1851 census, Herne Hill with parents and siblings; 1861, Herne Hill with Uncle James Horsburgh McDonald and siblings; 1871, Herne Hill occupation teacher, with Uncle James and brother John McDonald Lane; 1881 Wellington College, occupation assistant schoolmaster. After that I had no trace of him, but I have just found a record of his death and funeral in the Reading Mercury Sat 6 Dec 1884. (via findmypast). The account is very detailed and full of Victorian relish for sentimentality and funeral pomp.
He was the Bursar of Wellington College and died 29th Nov 1884, funeral Mon 1st Dec, Funeral procession 250 yards long, list of mourners, hymns sung at the service in Crowthorne, "Dead March" in Saul as coffin carried to the grave which was lined with green moss and white chrysanthemums and description of the coffin(polished oak with massive brass handles).
The interesting thing is that the leading mourners were "deceased's father and two nephews", the two nephews could have been Clayton Arbuthnot Lane who was 16, and his brother ,Frank McDonald Lane who was 14.
I checked the National Probate Calendar (ancestry) which has Charles Henry Lane, late of Wellington College, probate granted to James Lane of Torrington in the county of Devon , Gentleman, the Father and next of kin!
This suggests that James Lane was alive in 1884, but where he had been and why were his children living with Uncle James H McDonald?
This might be a massive red herring but all the evidence seems to fit, so good luck.
Ellen